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Qualification type
PhD / MPhil

Biological Sciences (Evolution, Ecology and Behaviour)

Funding
Self-funded
Study mode
Full-time
Part-time
Duration
3-4 years
5-6 years
Apply by
Year round
Start date
Year round
Subject area
Biological and Biomedical Sciences

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Overview

We study patterns in the natural environment, from the behaviour of individual organisms to dynamics of populations and the diversity of communities. Our aim is to go beyond description of these patterns - we wish to understand the ecological and evolutionary factors that drive them.

Introduction

Research in the Institute of Infection, Veterinary and Ecological Sciences spans the range of biosciences. From the fine-scale resolution of biomolecular structure, through the genome-wide analysis of cells and organisms, to the evolution and ecology of organisms in their natural environment. Underpinning this diversity, we offer world-class facilities and infrastructure for research, and superb links to internationally excellent partners in both the academic and industrial sectors.

Within this Institute, the Department of Evolution, Ecology and Behaviour works with viruses, bacteria, fungi, animals, insects, plants and humans, using state-of-the-art technologies and mathematical models to understand the principles that underlie biological diversity. The research seeks to answer fundamental questions and address key issues of contemporary concern.

Research topics

Ecology and evolution of infectious disease

We seek to understand the ecological and evolutionary dynamics of disease in wildlife populations. Our study systems are diverse, from microbes and their phage, through interactions in insects, wild rodent communities in the UK and abroad, to the microbes in the cystic fibrosis lung. To better understand the ecological and evolutionary behaviour of these systems, we employ field, genetic, genomic and mathematical approaches.

Evolution: phenotype to genome

Evolution has traditionally been studied at the level of the phenotype. With the emergence of new genomic technologies, we have an increasing ability to also examine how evolution is reflected in the genome, and to use techniques of molecular ecology to give insights into evolutionary processes.

Our current research is trying to understand the molecular basis of adaptation in the peppered moth, and in host-parasite interactions, and what these details tell us about the process of evolution. We are also employing molecular ecology tools to understand how geography and history have combined to structure current species.

Mammalian behaviour and evolution

We investigate mammalian behaviour, commonly working from molecules through to behaviour. We are particularly interested in:

  • Chemical communication
  • Reproductive strategies (with particular focus on mechanisms and evolutionary consequences of pre- and post-copulatory sexual selection)
  • Reproductive cooperation and conflict
  • The drivers of social structure and dispersion
  • Interactions between behaviour and disease.

We have an applied interest in the epidemiology of infections in domestic animals and in using ideas from evolution and behaviour to improve animal welfare.

Ecology: phenotype to ecosystem

We are interested in how individuals, species and communities respond to change, and how to mitigate risk.

We have particular interest in conservation biology, especially:

  • The metapopulation dynamics of threatened odonates
  • Thermal ecology, particularly effects of warming on organism size and ecological rates
  • Using protists as model organisms to study population dynamics, ecophysiology, distributional patterns, food web structure and the flow of energy and biomass through aquatic food webs
  • Phenotypic plasticity, especially maternal effects and developmental thresholds and their effects on population dynamics.
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How to apply

  1. 1. Identify potential supervisors

    It’s useful to contact potential supervisors before you begin your application. Explore our staff lists for the Institute of Infection, Veterinary and Ecological Sciences for details of potential supervisors.

  2. 2. Prepare your application documents

    You may need the following documents to complete your online application:

    • A research proposal (this should cover the research you’d like to undertake)
    • University transcripts and degree certificates to date
    • Passport details (international applicants only)
    • English language certificates (international applicants only)
    • A personal statement
    • A curriculum vitae (CV)
    • Contact details for two proposed supervisors
    • Names and contact details of two referees.
  3. 3. Apply

    Finally, register and apply online. You'll receive an email acknowledgment once you've submitted your application. We'll be in touch with further details about what happens next.

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My qualifications are from United Kingdom.

Fees and funding

Your tuition fees, funding your studies, and other costs to consider.

Tuition fees

UK fees (applies to Channel Islands, Isle of Man and Republic of Ireland)

Full-time place, per year - £5,006
Part-time place, per year - £2,503

International fees

Full-time place, per year - £31,250
Part-time place, per year - £15,650

Fees stated are for the 2025-26 academic year.


Additional costs

We understand that budgeting for your time at university is important, and we want to make sure you understand any costs that are not covered by your tuition fee. This could include buying a laptop, books, or stationery.

Find out more about the additional study costs that may apply to this degree.


Funding your PhD

If you're a UK national, or have settled status in the UK, you may be eligible to apply for a Postgraduate Doctoral Loan worth up to £30,301 to help with course fees and living costs.

There’s also a variety of alternative sources of funding. These include funded research opportunities and financial support from UK research councils, charities and trusts. Your supervisor may be able to help you secure funding.


Scholarships and bursaries

We offer a range of scholarships and bursaries that could help pay your tuition fees and living expenses.

Duncan Norman Research Scholarship

If you’re awarded this prestigious scholarship, you’ll receive significant funding to support your postgraduate research. This includes full payment of your PhD fees and a cash bursary of £17,000 per year while you study. One award is available in each academic year.

John Lennon Memorial Scholarship

If you’re a UK student, either born in or with strong family connections to Merseyside, you could be eligible to apply for financial support worth up to £12,000 per year for up to three years of full-time postgraduate research (or up to five years part-time pro-rata).

Sport Liverpool Performance Programme

Apply to receive tailored training support to enhance your sporting performance. Our athlete support package includes a range of benefits, from bespoke strength and conditioning training to physiotherapy sessions and one-to-one nutritional advice.

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Facilities and resources

The Institute of Infection, Veterinary and Ecological Sciences supports modern, well-equipped research laboratories on the University’s main Liverpool campus, including the Ronald Ross, Biosciences, Nuffield and IC2 Buildings. We also have facilities in Kenya, Malawi and Ethiopia.

Researchers benefit from access to world-class equipment and facilities via the Liverpool Shared Research Facilities, many of which are based within the Institute.

Virtual tour

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Meet our PhD researchers

These are the trailblazers, the leaders, the true originals. Our postgraduate researchers are shaping their future, and the future of the world we live in, advancing knowledge with each discovery.

Four postgraduate students walking on campus.

Chat with our students

Want to find out more about studying a PhD with us? Chat with our postgraduate research student ambassadors and ask any questions you have.

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Entry requirements

Discover the qualifications you'll need to apply for this course.

You’ll normally be expected to hold both:

  • A UK honours degree, or overseas equivalent, at 2:1 or above, in a relevant subject
  • A UK master’s degree, or overseas equivalent, in a relevant subject.


English language requirements

You'll need to demonstrate competence in the use of English language, unless you’re from a majority English speaking country.

We accept a variety of international language tests and country-specific qualifications.

International applicants who do not meet the minimum required standard of English language can complete one of our Pre-Sessional English courses to achieve the required level.

Qualification Details
IELTS 6.5 overall, with no component below 5.5
TOEFL iBT 88 overall with minimum scores of listening 17, writing 17, reading 17 and speaking 19. TOEFL Home Edition not accepted.
Pearson PTE Academic 61 overall, with no component below 59

Pre-sessional English

Do you need to complete a Pre-sessional English course to meet the English language requirements for this course?

The length of Pre-sessional English course you’ll need to take depends on your current level of English language ability.

Pre-sessional English in detail

If you don’t meet our English language requirements, we can use your most recent IELTS score, or the equivalent score in selected other English language tests, to determine the length of Pre-sessional English course you require.

Use the table below to check the course length you're likely to require for your current English language ability and see whether the course is available on campus or online.

Your most recent IELTS score Pre-sessional English course length On campus or online
6.0 overall, with no component below 5.5 6 weeks On campus
5.5 overall, with no component below 5.5 10 weeks On campus and online options available
5.5 overall, with no more than one component below 5.5, and no component below 5.0 12 weeks On campus and online options available
5.5 overall, with no component below 4.5 20 weeks On campus
5.0 overall, with no component below 4.5 30 weeks On campus
4.5 overall, with no more than one component below 4.5, and no component below 4.0 40 weeks On campus

If you’ve completed an alternative English language test to IELTS, we may be able to use this to assess your English language ability and determine the Pre-sessional English course length you require.

Please see our guide to Pre-sessional English entry requirements for IELTS 6.5 overall, with no component below 5.5, for further details.

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Contact us

Have a question about this research opportunity or studying a PhD with us? Please get in touch with us, using the contact details below, and we’ll be happy to assist you.

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